Alexander Vanin, arrested and convicted for espionage, hid a list of agents operating in the U.S. His control, Mishenko of the EEPR, has been ordered to get the list but Vanin refuses to give in because then Mishenko would cut him loose. The IMF must get the list.
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In London U.N.C.L.E. agent Brian Morton is planning on getting married, much to Waverly's disapproval. Solo and Illya help Morton on his current mission:stopping THRUSH agent Commander Kroler from stealing the plans for a new supersubmarine. Waverly tries to dissuade Laura Adams, Morton's fiancee, but to no avail. Morton and Laura are then kidnapped by Kroler, and Kroler threatens to kill Laura if Morton does not obtain the sub for him.
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Two cousins steal $500 million worth of jewels from a diamond syndicate and both U.N.C.L.E. and THRUSH want to find out how they pulled it off.