Lana receives startling news from Mary Eunice. Meanwhile, Arden sets out to learn what happened to Grace and Monsignor Howard takes on a true believer.
The team attempts to rally behind Captain Cragen after he comes under investigation for the death of the escort beside whose body he awoke. Unfortunately, they are hampered by an aggressive district attorney and a replacement captain whose management style differs greatly from Cragen's.
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The team investigates the death of an underage escort who died during a raucous bachelor party with VIP guests. The trail uncovers publicity-shy attendees, a fight for supremacy between two escort services and an old friend of the squad.
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As Meredith puts Zola to bed and falls asleep, she begins to wonder -- what if her mother had never had Alzheimer's and she'd had loving, supportive parents? The reverberations of a happy Meredith Grey change the world of Seattle Grace as we know it. What if she had never met Derek in that bar and he had never separated from Addison? What if Callie and Owen had become a couple long before she met Arizona? And what if Bailey never evolved from the meek intern she once was?
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