While Coulson's team infiltrate the ATCU base, Coulson interrogates Rosalind about what they discover during the break-in. Meanwhile, Gideon explains to Grant the true origins of Hydra.
While Fitz helps Jemma recover the information from her phone that could help the team open the portal, Coulson and Rosalind join forces to track down Andrew after Lincoln comes in out of the cold to confirm that Andrew is Lash.moreless
Coulson accompanies Rosalind to the ATCU facility, while Daisy, Mack, and Lance run surveillance on Banks. Meanwhile, Melinda takes Bobbi with her to track down the man who attacked Andrew.
Jemma tells Fitz about what happened to her on the alien planet... and the man who helped her survive.
While Coulson and Rosalind launch their first joint operation to find Lash, Melinda worries that Lance is too invested in his attempts to kill Grant.
While Mack and Daisy try to get to Lincoln before the ACTU does, Lance and Melinda try to get close to Grant. Meanwhile, Fitz nurses Jemma back to health after her rescue.
The team turns to Elliott for help figuring out where the monolith took Jemma, and he leads them to a castle in Gloucestershire. Meanwhile, Lance prepares to hunt down Grant and turns to Melinda for help.moreless
While Fitz tries to get Jemma back, Coulson and Skye discover that another organization is also hunting Inhumans.
Tensions rise between S.H.I.E.L.D. and the Inhumans, with Skye caught in the middle. Meanwhile, Coulson reveals what he's been doing with the missing millions.
Coulson and Gonzales are forced to join forces with an unreliable ally to defeat Hydra.
While Skye questions her future with the Inhumans, Melinda must decide her future in the war between the two sides of S.H.I.E.L.D.
While Skye meets a fellow "Inhuman," Coulson takes on Gonzales for the future of S.H.I.E.L.D.
Skye must make a decision about her powers, while Coulson finds himself going up against... S.H.I.E.L.D.
The team deals with the revelation of Skye's altered nature, while Bobbi and Mack tell Lance where their true allegiance lie. Meanwhile, Grant and Agent 33 go on a personal mission.
To expose S.H.I.E.L.D.'s supposed crimes to the world, Cal gathers a team of "super-villains" and lures Coulson back to his hometown. Meanwhile, Melinda brings in her ex-husband to help evaluate Skye's psychological condition after her recent transformation.moreless
After a battle, Lady Sif finds herself on Earth with amnesia, and turns to S.H.I.E.L.D. for help. Meanwhile, Bobbi and Mack continue their own operation within the team, risking exposure.
While Jemma secures the Kree city, Skye is put into quarantine. Meanwhile the team carries out an operation against Hydra to eliminate the people who arranged Trip's death.
Peggy races against time to stop Ivchenko and Dottie from carrying out Leviathan's master plan using Stark's stolen technology.
Peggy is cornered. Meanwhile, the SSR closes in on Howard Stark.
Daniel finds the final piece of the puzzle to tie Peggy to the nightclub and Brannis, while Peggy and Jarvis try to track down the woman who seduced Stark and gained access to his weapons vault.moreless
Peggy requests assignment to Jack's mission to Belarus and gets it when she provides the Howling Commandos as backup. Meanwhile, Roger reaches out to Jarvis, and Dottie pays a visit to Peggy's apartment.moreless
While Roger follows a lead to Europe, Stark turns up in Peggy's life again.
The SSR arrests Edwin after tracing Howard's car from the Roxxon explosion. Meanwhile, Peggy continues her pursuit of Leviathan.
After World War II is over, Peggy finds life as a glorified secretary for the SSR boring. When Howard Stark is framed for the sale of his weapons to the enemy, he asks Peggy to help him recover them and prove his innocence.moreless
Peggy pursues Brannis and the missing bombs, and Jarvis insists on helping her. Meanwhile, the SSR gets photos of the mystery blonde seen with Spider and speaks with Hugh Jones, the owner of the imploded factory.moreless
Coulson and his team prepare to destroy the Kree city, but learn where Whitehall and his people are drilling in and go to take them on. Meanwhile, Skye meets her father for the first time, and he reveals his plan to get revenge on Whitehall for killing Skye's mother.moreless
Coulson and half of the team head to San Juan to enter the Kree city, while the other half go to Vancouver to keep Raina from falling into Hydra's hands. However, Grant is one step ahead of them.moreless
While Coulson's team tries to track down the secret, they meet the Doctor. Meanwhile, Bobbi interrogates Sunil and Ward kidnaps his older brother.
Someone is killing people and leaving the alien writing etched on the corpses. As Coulson's team pursues the murderer, they discover a connection to their director's past.
Hydra impersonates S.H.I.E.L.D. and launches an attack on the UN to turn the world against Coulson and his people. As they rush to find the source of Hydra's new weapons, Coulson negotiates with the American Senator spearheaading the efforts to destroy S.H.I.E.L.D.: Grant's older brother Christian.moreless