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Cece deals with last-minute wedding catastrophes while Jess gets a new job opportunity. Nick bonds with Jess's dad, Schmidt navigates his new relationship with an old girlfriend, and Winston holds out hope that everyone hasn't forgotten his birthday.
In a series of flashbacks Jess, Nick, Schmidt, Winston and Cece tell one another how they each lost their virginity. Not surprisingly, when they realize the stories directly affect their present day sex lives, one relationship will change forever.