Origins brings viewers back to the beginning, before Nikita became a one-woman killing machine. On the streets and strung out on drugs, Nikita was well on her way to an early death or a long stint in prison. Carla is the one person who can help Nikita, whether Nikita likes it or not.
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Percy continues on this mission to take back Division by threatening to Carla, Nikita's former mentor who rescued Nikita from the streets when she was a junkie. Determined to reach Carla before Percy can get to her, Nikita uses Madeline to send Carla a message. However, Percy intercepts the message, and he and Nikita race to see who can get to Carla first. Meanwhile, Amanda tells Alex that the time has come for her to reclaim Zetrov and reveal her true identity to the world.
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In Clean Sweep, Percy holds members of Oversight hostage as leverage to make Amanda meet his demands. He threatens to let deadly gas loose inside Division. One of his hostages is Sean's mother, and Sean goes to Nikita seeking help. Nikita must decide whether to save the innocent lives within Division or let the enemy of her enemy become her friend.
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Desperate to protect his mother, Sean attacks Nikita and Michael. Meanwhile, Percy gains the upper hand on Amanda and threatens to take down Division unless she releases him.
Alex's mission to bring down Gogol comes to a head when she infiltrates her childhood home and faces ghosts from her past. Nikita, working on a separate mission, finds herself at the mansion as she tries to steal Semak's black box. When Nikita and Alex cross paths, they decide to team up, but the pair faces a major obstacle. Meanwhile, Michael and Owen join forces, hoping to uncover Percy's secret plan with the Guardians.
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With Michael out of town, Nikita looks to Owen for help as she tries to steal another black box from one of the Guardians. However, she learns that Owen is working for an unlikely employer that could jeopardize her whole mission. Amanda discovers that Alex received some confidential information about Semak and that Percy must have been the one to give her that information. Furious that Percy undermined her authority, Amanda strips him of all his amenities. Back in Russia, Alex finally makes her move on Semak.
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Oversight hosts an emergency meeting after realizing that Nikita is stealing their funding. Birkoff helps Nikita spy on the meeting, allowing her to identify the other members of Oversight, but things don't go as planned when Sean captures Birkoff. Amanda demands that Birkoff reveal where Nikita is hiding, but Birkoff refuses to give up her location, and Amanda tortures him. Meanwhile, Alex begins to execute her plan to reclaim her father's empire. With a price on her head, Alex decides that the the easiest way to get into Russia undetected is the same way she was smuggled out – through the illegal sex ring.
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Michael becomes concerned when he learns that Cassandra is in trouble, and he rushes off to protect her. Michael discovers that General Tupelov has accused her of stealing two hundred million dollars and has threatened to kill her unless she returns the money. However, some inconsistencies in Cassandra's story cause Nikita to wonder what Cassandra might be hiding. Meanwhile, Percy presents Alex with a very tempting offer that could provide her with the resources to take out Semak.
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From behind bars, Ryan uncovers patterns that reveal how Oversight gets its funding. He sets up a meeting with Nikita to fill her in on the details, but Amanda intercepts their coded messages and alerts Oversight, who sends in one of their members own to question Ryan. Nikita and Michael scramble to break Noah out of prison before Oversight reaches him. Meanwhile, Alex is being tailed by Gogol and comes face to face with someone from her past.
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While searching through the black box for information on Division, Birkoff comes across a copy of Nikita's birth certificate. Nikita and Michael locate Nikita's father, Richard, but the reunion does not go as Nikita had planned. Percy tells Alex that the black box in Nikita's possession contains information that could help her avenge her father's death.
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Michael learns that a woman he once romanced under Division orders, Cassandra, is in danger. Feeling guilty for how things ended, Michael wants to help Cassandra. Nikita, however, suspects that there might be more motivating Michael than his guilty conscience. Amanda manipulates Alex into helping Division on a mission, but Sean gets caught in Amanda's starts to wonder about Division's true motives.
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Nikita is shocked when she discovers that her old Division partner, Kelly, escaped from a Turkish prison. Nikita assumed that Kelly had died years ago and always felt responsible for her death, which occurred on the mission where Nikita escaped from Division. Knowing Amanda will hunt down after Kelly, Nikita tells Michael they have to travel to Turkey to help protect her. Amanda sends Alex and Roan to Europe to prevent Nikita and Kelly from reuniting.
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Nikita is upset when Birkoff tells her that information in the black box indicates a man named Ramon, who she captured years earlier and turned over to Percy, is now working for Division as an assassin. Amanda convinces Alex to kill Anton Kochenko, a man who once worked for her father, and sets up Ramon so that he will be charged with the murder. However, when Nikita interferes with Amanda's plan, Amanda retaliates by taking what Nikita values most.
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Nikita and Michael investigate an old corrupt Division program called P9 that appears to have resurfaced. During their investigation, they run into Owen, who informs them that he is searching for Dr. Joseph Mars, the man who not only masterminded P9, but also invented the Regimen and could help Owen wean himself off the dangerous drug. Michael is reluctant to trust Owen again, but Nikita decides that the three should join forces. Back at Division, Percy demands to see Alex.
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Nikita and Michael plan to take down Division and Oversight, using the contents of the black box to help them with their mission. Amanda is now leading Division, but Oversight has assigned one of their agents, Sean, to keep a a close watch on her. Amanda's first mission is to retrieve the black box from Nikita, and she pursues Alex's help in the mission. Alex refuses to cooperate until Amanda tells her she will help her find the man who killed Alex's father. Meanwhile, Nikita and Michael receive help from an unlikely ally during their mission.
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