Formerly prisoners of the Breen, Worf and Ezri Dax are now prisoners of the Dominion, aboard a Jem'Hadar vessel. Also on board is Weyoun and the Female Changeling, forming an alliance with the Breen against the Federation. The Female Changeling has to concentrate hard in order to maintain her normal, smooth-skinned humanoid form. Legate Damar voices his displeasure at the new alliance, as he, as the Cardassian leader, was not informed and left out of the negotiations with the Breen before now. Also, the Cardassians must cede a few planets to the Breen seemingly without any return. Damar becomes even more angry, when he learns that Septimus III has fallen to the enemy, even though Weyoun had promised to "deal" with the situation.
General Martok and Captain Sisko are discussing the war, and their conversation drifts toward marriage, and the resemblance of marriage to war.
Dukat, still posing as Bajoran farmer Anjohl Tennan, continues his manipulation of Kai Winn Adami. He continually points to her "prophecy", and how it warned her of the Emissary of the Prophets straying from course.
Kasidy finds it difficult being to married to the Emissary and politely declines Benjamin's request that she perform a Bajoran fertility ceremony. Meanwhile, Worf and Ezri continue to plot their escape, now from the clutches of the Jem'Hadar. They soon turn to arguing, invoking the name of Jadzia, Dax's previous host who was married to Worf.
When Damar announced to them that they will be tried as war criminals and executed, Weyoun offers them a chance to escape death, by supplying him with information. When he insults Ezri by mentioning her "relationship" with Julian Bashir, Worf immediately snaps Weyoun's neck, killing him instantly. Damar laughs heartily, happy to see his tormentor dead, despite knowing that he would soon be replaced by the next clone.
The Dominion send a brand new Weyoun who allows the Breen to examine their military database, much to Damar's chagrin. Even worse, Damar is demoted, told by Weyoun that he now reports to the Breen. Having fashioned a club, Worf and Ezri break out of their cell, only to be chased down in the corridors, recaptured. Their absence on Deep Space 9 is noticed by everyone, especially Quark, who continues to pour (the absent) Ezri her usual drink at the same time every day. The mood is dreary in Quark's, but the moment seems to be especially poignant to Bashir.
Later, Kai Winn experiences her second vision from the "Prophets," who reveal to her that they are indeed the Pah-wraiths, not the celestial Prophets. She consults with the Orb of Prophecy, but the Prophets do not speak to her. Dukat reveals that he is a servant of the Pah-wraiths, and that he has been brought to the Kai for a purpose: to unite against the Emissary and the Prophets of the Celestial Temple. She meets with Kira, and bares her soul to her. Kira advises her to step down as Kai, because it was power that led Winn away from the Prophets. However, Winn is unwilling to give up the political power that her position has given her.
With the Breen suddenly replacing the Cardassians as the Dominion's best Alpha Quadrant ally, Damar feels like the Cardassians are being squeezed out of control. Therefore, he boldly shoots the Jem'Hadar guards accompanying Worf and Ezri, allowing their escape. Worf and Ezri, who had come to a truce and agreed to remain friends, are given a message from Damar to the Federation: that he will be their sole ally on Cardassia.
Kai Winn, feeling that the Prophets have turned their back on her, pledges her allegiance to the Pah-wraiths. She and Dukat vow to destroy any who would stand in their way, including the Federation, the Prophets, and the Emissary.