Bath time is interrupted by an invasive insect hopping from one set of skimpy skivvies to another. The Strike Witches are up in arms over this shocking breach of their britches, and they'll do whatever it takes to protect their panties!
A menacing new breed of Neuroi looms high above the clouds, and the Strike Witches must equip themselves with experimental rocket boosters in order to reach the unprecedented altitude required to destroy their enemy!
When Sakamoto's strike unit malfunctions, Yoshika is forced into a solo dogfight against a rapidly-regenerating Neuroi! The bombastic beauties of the 501st Joint Fighter Wing race to provide backup, but will they arrive in time?
Yoshika's plans to become a doctor are put on hold after a mysterious letter from her father leads to a reunion with Sakamoto. Elsewhere, her former Strike Witch comrades take to the air to battle a new breed of Neuroi!
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