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Illya parachutes out of a plane carrying THRUSH code documents, but lands in the desert at an abandoned Foreign Legion post run by Captain Basil Calhoun. Illya is accompanied by a stewardess, Barbara while Solo races to find them before THRUSH does.
A top-level U.N.C.L.E. conference is to be held in Switzerland, but a nearby boys' school is actually a Thrush front run by Mother Fear and Dennis Jenks that is training its students to be assassins. Solo is captured by the boys, and, when he refuses to reveal the conference location, is forced to operate the controls of two electric trains so they do not collide with each carrying a vial of deadly nerve gas. Illya and Anna Paola, a social worker who resent children, are also captured. The escape, and arrive at the conference just as the boy's choir is ready to kill Waverly and the others with Thrush rifles from under their robes.moreless
With the aid of Dr. Elias Swan, THRUSH has developed a strain of deadly bees. THRUSH agent Mr. Mozart captures Illya and Tavia Sandor and uses a high-frequency sound machine that threatens to shatter their eardrums to force Illya to take him to U.N.C.L.E. headquarters so he can release the bees. Illya does so, but Mozart is intercepted, and in a battle on a rooftop, is shot and the bees get loose.moreless
Illya infiltrates a South American prison camp where THRUSH is storing the "Ultimate Computer."
Thrush is so eager to get their hands on a boy genius that they murder the boy's father. But then they have to deal with the boy's madcap aunt.
Thrush Eastern in Japan acquires a volcano-activating device, and Solo and Illya, along with Cricket Okasada, a film student, infiltrate a toy store and a karate school to find it. Thrush leader Mr. Kutuzov oversees local Thrush head Harada in the operation, and Solo finds himself fighting for his life against a life-size sword-wielding puppet.moreless
Illya goes to find Thrush's vaporizing machine, but is captured by Sophie, the headstrong daughter of the local tribal chief, Sulador. In New York, Solo convinces retiring Thrush agent David Lewin that Thrush intends to kill him, and elicits his help. Lewin and Solo are taken to the base in the desert, just as Illya leads the band of nomads, whose confidence he has gained, in an attack on the base.moreless
Napoleon is given the job of searching for Thrush security records.
U.N.C.L.E. and Thrush agents compete to gain possession of a mind-reading device created by a magician who has been murdered by Thrush.
The "Re-collectors" claim to be trying to locate works of art stolen by the Nazis but all is not as it appears.
Mr. Alexander, a megalomaniac industrialist, plans to conquer the world like his namesake, Alexander the Great. After escaping the pit and the pendulum in the tomb, Solo battles a bodybuilder in a gymnasium, while at Alexander's farm in Virginia, Illya is nearly cut in two by various farm implements. He is then wrapped in bandages from head to toe in preparation for being mummified by Prof. Kavon. The UNCLE agents seek to stop Alexander from assassinating the president of an Asian country.moreless
Alexander, a megalomaniac industrialist, plans to conquer the world like his namesake, Alexander the Great, by breaking each of the Ten Commandments along the way. He has his henchman Parviz steal the army's "will gas" to help him do so, and Solo investigates. Solo encounters Tracey Alexander, Alexander's ex-wife, who wants to tag along with Solo to get the money Alexander owes her. The trail leads Solo, Illya, and Tracey to a posh party at Alexander's, where Solo defeats their host in a human chess game; to a tomb, where Alexander and his aide Dr. Kavon leave Solo tied under a descending scimitar while Illya and Tracey are hanging over a bottomless pit with a candle burning their rope; to Alexander's farm in Virginia, where Illya is nearly cut in two by various farm implements.moreless